OK, I have to ask this since it seems obvious.
If I do not have the right voltage right out of the input / power transformer, why hasn't anyone said, THAT looks like your problem and suggested I replace it ? It seems like you have me checking voltages with and without tubes, removing caps and diodes to troubleshoot incorrect voltage and voltage drops, but what we are starting with is already too low. If it was good at the start, but but gets pulled down when I plug it into the board, I would understand looking for issues down stream, but ..."Volts across the red wires, unplugged from board = 168 VAC " tells me that it's bad even before I plug power into the board. Yes, the green pair starts good and drops, but isn't the transformer already suspect with the high volt input being down ? If I were to put a 100ohm resistor across the green wires, if that section of the transformer were healthy, I should see 6.3vac on the resistor.